Year Published

Displaying 1-100 of 114 Publications


Damian-Serrano, A., E.D. Hetherington, C.A. Choy, S.H.D. Haddock, A. Lapides, and C.W. Dunn. 2022. Characterizing the secret diets of siphonophores (Cnidaria: Hydrozoa) using DNA metabarcoding. PLOS ONE, 17(e0267761): 1–24.

Hetherington, E.D., A. Damian-Serrano, S.H.D. Haddock, C.W. Dunn, and C.A. Choy. 2022. Integrating siphonophores into marine food-web ecology. Limnology and Oceanography Letters, 7: 81–95.

Hetherington, E.D., C.A. Choy,  E.V. Thuesen, and S.H.D. Haddock. 2022. Three distinct views of deep pelagic community composition based on complementary sampling approaches. Frontiers in Marine Science, 9(864004): 1–10.

Johnson, S.B.,  J.R. Winnikoff,  D.T. Schultz,  L.M. Christianson,  W.L. Patry, C.E. Mills, and S.H.D. Haddock. 2022. Speciation of pelagic zooplankton: Invisible boundaries can drive isolation of oceanic ctenophores. Frontiers in Genetics, 13(970314): 1–17.

Matsumoto, G.I., L.M. Christianson, B.H. Robison, S.H.D. Haddock, and S.B. Johnson. 2022. Atolla reynoldsi sp. nov. (Cnidaria, Scyphozoa, Coronatae, Atollidae): A new species of coronate Scyphozoan found in the Eastern North Pacific Ocean. Animals, 12: 1–21.


Christianson, L.M., S.B. Johnson, D.T. Schultz, and S.H.D. Haddock. 2021. Hidden diversity of Ctenophora revealed by new mitochondrial COI primers and sequences. Molecular Ecology Resources, 22: 283–294.

Damian-Serrano, A., S.H.D. Haddock, and C.W. Dunn. 2021. The evolution of siphonophore tentilla for specialized prey capture in the open ocean. Proceedings of the National Academy of Sciences, 118: 1–9.

Elor, A., T. Thang, B.P. Hughes, A. Crosby, A. Phung, E. Gonzalez, K. Katija, S.H.D. Haddock, B.E. Erwin, E.J. Martin, and L. Takayama. 2021. Catching jellies in immersive virtual reality: A comparative teleoperation study of ROVs in underwater capture tasks. Proceedings of the 27th ACM Symposium on Virtual Reality Software and Technology, VRST '21 (17): 1–10.

Helm, R.R., N. Clark, H. Harden-Davies, D. Amon, P. Girguis, C. Bordehore, S. Earle, M.J. Gibbons, Y. Golbuu, S.H.D. Haddock, J.D.R. Houghton, J. Javidpour, D.J. McCauley, L. Morgan, D. Obura, E.A. Pakhomov, K.A. Pitt, J. Jimenez Ramon, R. Sumaila, and J.B. Thiebot. 2021. Protect high seas biodiversity. Science, 372: 1048–1049.

Meech, M.E., C.E. Mills, S.H.D. Haddock, and R.W. Meech. 2021. Two swimming modes in Trachymedusae; bell kinematics and the role of giant axons. Journal of Experimental Biology, 224: 1–12.

Miller, S.D., S.H.D. Haddock, W.C. Straka III, C.J. Seaman, C.L. Combs, M. Wang, W. Shi, and S. Nam. 2021. Honing in on bioluminescent milky seas from space. Scientific Reports, 11: 1–10.

Puente-Tapia, F.A., R. Gasca, A. Schiariti, and S.H.D. Haddock. 2021. An updated checklist of ctenophores (Ctenophora: Nuda and Tentaculata) of Mexican seas. Regional Studies in Marine Science, 41: 1–20.

Schultz, D.T., W.R. Francis, J.D. McBroome, L.M. Christianson, S.H.D. Haddock, and R.E. Green. 2021. A chromosome-scale genome assembly and karyotype of the ctenophore Hormiphora californensis. G3 Genes|Genomes|Genetics, 11(11): 1–12.

Damian-Serrano, A., S.H.D. Haddock, and C.W. Dunn. 2021. The evolutionary history of siphonophore tentilla: Novelties, convergence, and integration. Integrative Organismal Biology, 3: 1–21.

Stefanoudis, P. V., L.M. Biancani, S. Cambronero-Solano, M.R. Clark, J.T. Copley, E. Easton, F. Elmer, S.H.D. Haddock, S. Herrera, I.S. Iglesias, A.M. Quattrini, J. Sigwart, C. Yesson, and A.G. Glover. 2021. Moving conferences online: lessons learned from an international virtual meeting. Proceedings of the Royal Society B, 288: 1–7.

Winnikoff, J.R., S.H.D. Haddock, and I. Budin. 2021. Depth- and temperature-specific fatty acid adaptations in ctenophores from extreme habitats. Journal of Experimental Biology, 224: 1–10.


Bessho-Uehara, M., W. Huang, W.L. Patry, W. E. Browne, J.K. Weng, and S.H.D. Haddock. 2020. Evidence for de novo biosynthesis of the luminous substrate coelenterazine in ctenophores. iScience, 23: 1–11.

Bessho-Uehara, M., W. R. Francis, and S. Haddock. 2020. Biochemical characterization of diverse deep-sea anthozoan bioluminescence systems. Marine Biology, 167: 1–19.

Drazen, J.C., C.R. Smith, K.M. Gjerde, S.H.D. Haddock, G.S. Carter, C.A. Choy, M.R. Clark, P. Dutrieux, E. Goetze, C. Hauton, M. Hatta, J.A. Koslow, A.B. Leitner, A. Pacini, J.N. Perelman, T. Peacock, T.T. Sutton, L. Watling, and H. Yamamoto. 2020. Midwater ecosystems must be considered when evaluating environmental risks of deep-sea mining. Proceedings of the National Academy of Sciences, 117(30): 17455–17460.

Leonardi, N.D., E.V. Thuesen, and S.H.D. Haddock. 2020. A sticky thicket of glue cells: A comparative morphometric analysis of colloblasts in 20 species of comb jelly (phylum Ctenophora). Ciencias Marinas, 46: 211–225.

Martini, S, D.T. Schultz, L. Lundsten, and S.H.D. Haddock. 2020. Bioluminescence in an undescribed species of carnivorous sponge (Cladorhizidae) from the deep sea. Science, 7: 1–19.

Schultz, D.T., J.M. Eizenga, R.B. Corbett-Detig, W.R. Francis, L.M. Christianson, and S.H.D. Haddock. 2020. Conserved novel ORFs in the mitochondrial genome of the ctenophore Beroe forskalii. PeerJ, 8: e8356.


Martini, S., L. Kuhnz, J. Mallefet, and S.H.D. Haddock. 2019. Distribution and quantification of bioluminescence as an ecological trait in the deep sea benthos. Scientific Reports, 9(14654).

Messié, M., I. Shulman, S. Martini, and S. Haddock. 2019. Using fluorescence and bioluminescence sensors to characterize auto- and heterotrophic plankton communities. Progress in Oceanography, 171: 76–92.

Winnikoff, J.R., W.R. Francis, E.V. Thuesen, and S.H.D. Haddock. 2019. Combing transcriptomes for secrets of deep-sea survival: Environmental diversity drives patterns of protein evolution. Integrative and Comparative Biology, 59(4): 786–798. PDF.


Babonis, L., M.B. Debiasse, W.R. Francis, L.M. Christianson, S.H.D. Haddock, M.Q. Martindale, and J.F. Ryan. 2018. Integrating embryonic development and evolutionary history to characterize tentacle-specific cell types in a ctenophore. Molecular Biology and Evolution, 35: 2940–2956.

Munro, C., S. Siebert, F. Zapata, M. Howison, A. Damian-Serrano, S.H. Church, F.E. Goetz, P.R. Pugh, S.H.D. Haddock, and C.W. Dunn. 2018. Improved phylogenetic resolution within Siphonophora (Cnidaria) with implications for trait evolution. Molecular Phylogenetics and Evolution, 127: 823–833.

Pugh, P.R., C.W. Dunn, and S.H.D. Haddock. 2018. A description of Tottonophyes enigmatica gen. nov., sp. nov. in a new family of calycophoran siphonophores (Hydrozoa), with notes on calycophoran canals. Zootaxa, 4415: 452–472. PDF.

Schultz, D.T., A.A. Kotlobay, R. Ziganshin, A. Bannikov, N.M. Markina, T.V. Chepurnyh, E.S. Shakhova, K. Palkina, S.H.D. Haddock, I.V. Yampolsky, and Y. Oba. 2018. Luciferase of the Japanese syllid polychaete Odontosyllis umdecimdonta. Biochemical and Biophysical Research Communications, 502: 318–323.


Choy, C.A., S.H.D. Haddock, and B.H. Robison. 2017. Deep pelagic food web structure as revealed by in situ feeding observations. Proc Biol Sci, 284: 1–10.

Francis, W.R., L.M. Christianson, and S.H.D. Haddock. 2017. Symplectin evolved from multiple duplications in bioluminescent squid. PeerJ, 5: e3633.

Haddock, S.H.D., L.M. Christianson, W.R. Francis, S. Martini, C.W. Dunn, P.R. Pugh, C.E. Mills, K.J. Osborn, B.A. Seibel, C.A. Choy, C.E. Schnitzler, G.I. Matsumoto, M. Messié, D.T. Schultz, J.R. Winnikoff, M.L. Powers, R. Gasca, W.E. Browne, S. Johnsen, K.L. Schlining, S. von Thun, B.E. Erwin, J.F. Ryan, and E.V. Thuesen. 2017. Insights into the biodiversity, behavior, and bioluminescence of deep-sea organisms using molecular and maritime technology. Oceanography, 30: 38–47. PDF.

Hoving, H.J., and S.H. Haddock. 2017. The giant deep-sea octopus Haliphron atlanticus forages on gelatinous fauna. Scientific Reports, 7: 1–4.

Hoving, H.J.T., S.L. Bush, S.H.D. Haddock, and B.H. Robison. 2017. Bathyal feasting: post-spawning squid as a source of carbon for deep-sea benthic communities. Proceedings of the Royal Society B, 284: 1–5.

Martini, S., and S.H.D. Haddock. 2017. Quantification of bioluminescence from the surface to the deep sea demonstrates its predominance as an ecological trait. Scientific Reports, 7: 1–11.

Winnikoff, J.R., T.M. Wilson, E.V. Thuesen, and S.H.D. Haddock. 2017. Enzymes feel the squeeze: biochemical adaptation to pressure in the deep sea. The Biochemist, 39: 26–29. PDF.


Fossette, S., K. Katija, J.A. Goldbogen, S. Bograd, W. Patry, M.J. Howard, T. Knowles, S.H.D. Haddock, L. Bedell, E.L. Hazen, B.H. Robison, T.A. Mooney, K.A. Shorter, T. Bastian, and A.C. Gleiss. 2016. How to tag a jellyfish? A methodological review and guidelines to successful jellyfish tagging. Journal of Plankton Research, 38: 1347–1363.

Francis, W.R., L.M. Christianson, M.L. Powers, C.E. Schnitzler, and S.H.D. Haddock. 2016. Non-excitable fluorescent protein orthologs found in ctenophores. BMC Evolutionary Biology, 16: 167.

Gasca, R., and S.H. Haddock. 2016. The rare deep-living hyperiid amphipod Megalanceoloides remipes (Barnard, 1932): Complementary description and symbiosis. Zootaxa, 4178: 138–144.

Pugh, P.R., and S.H.D. Haddock. 2016. A description of two new species of the genus Erenna (Siphonophora, Physonectae, Erennidae), with notes on recently collected specimens of other Erenna species. Zootaxa, 4189: 401–446.


Francis, W.R., N.C. Shaner, L.M. Christianson, M.L. Powers, and S.H.D. Haddock. 2015. Occurrence of isopenicillin-N-synthase homologs in bioluminescent ctenophores and implications for coelenterazine biosynthesis. PLoS One, 10: e0128742. PDF.

Gasca, R., E. Suárez-Morales, and S.H.D. Haddock. 2015. Sapphirina iris Dana, 1849 and S. sinuicauda Brady, 1883 (Copepoda, Cyclopoida): Predators of salps in Monterey Bay and the Gulf of California. Crustaceana, 88: 689–699.

Haddock, S.H.D., and C.W. Dunn. 2015. Fluorescent proteins function as a prey attractant: Experimental evidence from the hydromedusa Olindias formosus and other marine organisms. Biology Open, 4(9): 1094–1104. Supplemental video.

Zapata, F., F.E. Goetz, S.A. Smith, M. Howison, S. Siebert, S.H. Church, S.M. Sanders, C.L. Ames, C.S. McFadden, S.C. France, M. Daly, A.G. Collins, S.H.D. Haddock, C.W. Dunn, and P. Cartwright. 2015. Phylogenomic analyses support traditional relationships within Cnidaria. PLoS One, 10: e0139068.


Bracken-Grissom, H., A.G. Collins, T. Collins, K. Crandall, D. Distel, C.W. Dunn, G. Giribet, S.H.D. Haddock, N. Knowlton, M.Q. Martindale, M. Medina, C.G. Messing, S. O'Brien, G. Paulay, N. Putnam, T. Ravasi, G. Rouse, J. Ryan, A. Schulze, G. Wörheide, M. Adamska, X. Bailly, J. Breinholt, W.E. Browne, C. Diaz, N. Evans, J.F. Flot, N. Fogarty, M. Johnston, B. Kamel, A.Y. Kawahara, T. Laberge, D. Lavrov, F. Michonneau, L.L. Moroz, T. Oakley, K.J. Osborne, S. Pomponi, A. Rhodes, M. Rodriguez-Lanetty, S.R. Santos, N. Satoh, R.W. Thacker, Y. Van de Peer, C.R. Voolstra, D.M. Welch, J. Winston, and X. Zhou. 2014. The Global Invertebrate Genomics Alliance (GIGA): Developing community resources to study diverse invertebrate genomes. Journal of Heredity, 105: 1–18. PDF.

Cazenave, F., C. Kecy, M. Risi, and S.H.D. Haddock. 2014. SeeStar: A low-cost, modular and open-source camera system for subsea observations. In Proceedings of Marine Technology Society/Institute for Electrical and Electronics Engineers Oceans 2014, St. John's, NL, Canada, 14–19 September 2014.

Francis, W.R., M.L. Powers, and S.H.D. Haddock. 2014. Characterization of an anthraquinone fluor from the bioluminescent, pelagic polychaete Tomopteris. Luminescence, 29: 1135–1140. PDF.

Gasca, R., R. Hoover, and S.H.D. Haddock. 2014. New symbiotic associations of hyperiid amphipods (Peracarida) with gelatinous zooplankton in deep waters off California. Journal of the Marine Biological Association of the United Kingdom, 95: 503–511.

Wilson, G., D.A. Aruliah, C.T. Brown, N.P.C. Hong, M. Davis, R.T. Guy., S.H.D. Haddock, K.D. Huff, I.M. Mitchell, M.D. Plumbley, B. Waugh, E.P. White, and P. Wilson. 2014. Best practices for scientific computing. PLoS Biology, 12: e1001745.


Francis, W.R., L.M. Christianson, R. Kiko, M.L. Powers, N.C. Shaner, and S.H.D. Haddock. 2013. A comparison across non-model animals suggests an optimal sequencing depth for de novo transcriptome assembly. BMC Genomics, 14: 167.

Hurt, C., S.H.D. Haddock, and W.E. Browne. 2013. Molecular phylogenetic evidence for the reorganization of the Hyperiid amphipods, a diverse group of pelagic crustaceans. Molecular Phylogenetics and Evolution, 67: 28–37.

Madin, L.P., W.M. Hamner, S.H.D. Haddock, and G.I. Matsumoto. Scuba diving in blue water: A window on ecology and evolution in the epipelagic ocean. In Research and Discovery: The Revolution of Science through Scuba, edited by Lang, M.A., R.L. Marinelli, S.J. Roberts, and P.R. Taylor, 71–82. Washington, D.C., Smithsonian Institution Scholarly Press: 2013.

Ryan, J.F., K. Pang, C.E. Schnitzler, A.D. Nguyen, R.T. Moreland, D.K. Simmons, B.J. Koch, W.R. Francis, P. Havlak, NISC Comparative Sequencing Program, S.A. Smith, N.H. Putnam, S.H.D. Haddock, C.W. Dunn, T.G. Wolfsberg, J.C. Mullikin, M.Q. Martindale, and A.D. Baxevanis. 2013. The genome of the ctenophore Mnemiopsis leidyi and its implications for cell type evolution. Science, 342: 1336–1344.

Siebert, S., P.R. Pugh, S.H.D. Haddock, and C.W. Dunn. 2013. Re-evaluation of characters in Apolemiidae (Siphonophora), with description of two new species from Monterey Bay, California. Zootaxa, 3702: 201–232.

Thuesen, E.V., and S.H.D. Haddock. 2013. Archeterokrohnia docrickettsae (Chaetognatha: Phragmophora: Heterokrohniidae), a new species of deep-sea arrow worm from the Gulf of California. Zootaxa, 3717: 320–328.


Condon, R.H., C.M. Duarte, K.A. Pitt, K.L. Robinson, C.H. Lucas, K.R. Sutherland, H.W. Mianzan, M. Bogeberg, J.E. Purcell, M.B. Decker, S.I. Uye, L.P. Madin, R.D. Brodeur, S.H.D. Haddock, A. Malej, G.D. Parry, E. Eriksen, J. Quiñones, M. Acha, M. Harvey, J.M. Arthur, and W.M. Graham. 2012. Recurrent jellyfish blooms are a consequence of global oscillations. Proceedings of the National Academy of Sciences of the United States of America, 110: 1000–1005.

Condon, R.H., W.M. Graham, C.M. Duarte, K.A. Pitt, C.H. Lucas, S.H.D. Haddock, K.R. Sutherland, K.L. Robinson, M.N. Dawson, M.B. Decker, C.E. Mills, J.E. Purcell, A. Malej, H. Mianzan, S.I. Uye, S. Gelcich, and L.P. Madin. 2012. Questioning the rise of gelatinous zooplankton in the world's oceans. BioScience, 62: 160–169.

Johnsen, S., T.M. Frank, S.H.D. Haddock, E.A. Widder, and C.G. Messing. 2012. Light and vision in the deep-sea benthos: I. Bioluminescence at 500–1000 m depth in the Bahamian Islands. Journal of Experimental Biology, 215: 3335–3343.

Powers, M.L., A.G. McDermott, N. Shaner, and S.H.D. Haddock. 2012. Expression and characterization of the calcium-activated photoprotein from the ctenophore Bathocyroe fosteri: Insights into light-sensitive photoproteins. Biochemical and Biophysical Research Communications, 431(2): 360–366.

Schnitzler, C.E., K. Pang, M.L. Powers, A.M. Reitzel, J.F. Ryan, D. Simmons, T. Tada, M. Park, J. Gupta, S.Y. Brooks, R.W. Blakesley, S. Yokoyama, S.H.D. Haddock, M.Q. Martindale, and A.D. Baxevanis. 2012. Genomic organization, evolution, and expression of photoprotein and opsin genes in Mnemiopsis leidyi: a new view of ctenophore photocytes. BMC Biology, 10: 1–26.

Shulman, I., B. Penta, M.A. Moline, S.H.D. Haddock, S. Anderson, M.J. Oliver, and P. Sakalaukus. 2012. Can vertical migrations of dinoflagellates explain observed bioluminescence patterns during an upwelling event in Monterey Bay, California?. Journal of Geophysical Research, 117: C01016.


Haddock, S.H.D., and C.W. Dunn. Practical computing for biologists. Sunderland, Massachusetts: Sinauer Associates, 2011.

Shulman, I., M.A. Moline, B. Penta, S. Anderson, M. Oliver, and S.H.D. Haddock. 2011. Observed and modeled bio-optical, bioluminescent, and physical properties during a coastal upwelling event in Monterey Bay, California. Journal of Geophysical Research-Oceans, 116: 1–13.


Bayha, K.M., M.N. Dawson, A.G. Collins, M.S. Barbeitos, and S.H.D. Haddock. 2010. Evolutionary relationships among scyphozoan jellyfish families based on complete taxon sampling and phylogenetic analyses of 18S and 28S ribosomal DNA. Integrative and Comparative Biology, 50: 436–455.

Haddock, S.H.D. 2010. A plea for meaningful colors in ODV. Limnology and Oceanography Bulletin, 19: 44–45.

Haddock, S.H.D., N. Mastroianni, and L.M. Christianson. 2010. A photoactivatable green-fluorescent protein from the phylum Ctenophora. Proceedings of the Royal Society B, 277: 1155–1160.

Pugh, P.R., and S.H.D. Haddock. 2010. Three new species of resomiid siphonophores (Siphonophora, Physonectae). Journal of the Marine Biological Association of the United Kingdom, 90: 1119–1143.

Thuesen, E.V., F.E. Goetz, and S.H.D. Haddock. 2010. Bioluminescent organs of two deep-sea arrow worms, Eukrohnia fowleri and Caecosagitta macrocephala, with further observations on bioluminescence in chaetognaths. Biological Bulletin, 219: 100–111.


Gorokhova, E., M. Lehtiniemi, S. Viitasalo-Frösen, and S.H.D. Haddock. 2009. Molecular evidence for the occurrence of ctenophore Mertensia ovum in the northern Baltic Sea and implications for current distribution of the invasive species Mnemiopsis leidyi. Limnology and Oceanography, 54: 2025–2033.

Moline, M.A., S.M. Blackwell, J.F. Case, S.H.D. Haddock, C.M. Herren, C.M. Orrico, and E. Terrill. 2009. Bioluminescence to reveal structure and interaction of coastal planktonic communities. Deep-Sea Research Part II, 56: 232–245.


Collins, A.G., B. Bentlage, A. Lindner, D. Lindsay, S.H.D. Haddock, G. Jarms, J.L. Norenburg, T. Jankowski, and P. Cartwright. 2008. Phylogenetics of Trachylina (Cnidaria: Hydrozoa) with new insights on the evolution of some problematical taxa. Journal of the Marine Biological Association of the United Kingdom, 88: 1673–1685.

Dunn, C.W., A. Hejnol, D.Q. Matus, K. Pang, W.E. Browne, S.A. Smith, E. Seaver, G.W. Rouse, M. Obst, G.D. Edgecombe, M.V. Sorensen, S.H.D. Haddock, A. Schmidt-Rhaesa, A. Okusu, R.M. Kristensen, W.C. Wheeler, M.Q. Martindale, and G. Giribet. 2008. Broad phylogenomic sampling improves resolution of the animal tree of life. Nature, 452: 745–749.

Haddock, S. H. D. 2008. Reconsidering evidence for potential climate-related increases in jellyfish. Limnology and Oceanography, 53: 2759–2762.

Schnitzler, C.E., R.J. Keenan, R. McCord, A. Matysik, L.M. Christianson, and S.H.D. Haddock. 2008. Spectral diversity of fluorescent proteins from the anthozoan Corynactis californica. Marine Biotechnology, 10: 328–342.

Staaf, D.J., S. Camarillo-Coop, S.H.D. Haddock, A.C. Nyack, J. Payne, R. Ramirez-Rojo, C.A. Salinas-Zavala, B.A. Seibel, L. Trueblood, C. Widmer, and W.F. Gilly. 2008. Natural egg mass deposition by the Humboldt squid (Dosidicus gigas) in the Gulf of California and characteristics of hatchlings and paralarvae . Journal of the Marine Biological Association of the United Kingdom, 88: 759–770.


Browne, W.E., S.H.D. Haddock, and M.Q.Martindale. 2007. Phylogenetic analysis of lineage relationships among hyperiid amphipods as revealed by examination of the mitochondrial gene, cytochrome oxidase I (COI). Integrative and Comparative Biology, 47: 815–830.

Mills, C.E., and S.H.D. Haddock. 2007. Key to the ctenophora. Light and Smith's Manual: Intertidal invertebrates of the central California coast: 189–199.

Mills, C.E., S.H.D. Haddock, C.W. Dunn, and P.R. Pugh. 2007. Key to the siphonophora. Light and Smith's Manual: Intertidal invertebrates of the central California coast: 150–166.

Sweeney, A.M., S.H.D. Haddock, and S. Johnsen. 2007. Comparative visual acuity of coleolid cephalopods. Integrative and Comparative Biology, 47: 808–814.


Collins, A.G., B. Bentlage, G.I. Matsumoto, S.H.D. Haddock, K.J. Osborn, and B. Schierwater. 2006. Solution to the phylogenetic enigma of Tetraplatia, a worm-shaped cnidarian. Biological Letters, 2: 120–124.

Gasca, R., E. Suárez-Morales, and S.H.D. Haddock. 2006. Symbiotic associations between crustaceans and gelatinous zooplankton in deep and surface waters off California. Marine Biology, 151: 233–242.

Haddock, S.H.D. 2006. Luminous marine organisms. Photoproteins in Bioanalysis: 25–47.

Miller, S.D, S.H.D. Haddock, C.D. Elvidge, and T.F. Lee. 2006. Twenty thousand leagues over the seas: The first satellite perspective on bioluminescent ‘milky seas’. International Journal of Remote Sensing, 27: 5131–5143.


Dunn, C.W., P.R. Pugh, and S.H.D. Haddock. 2005. Marrus claudanielis, a new species of deep-sea physonect siphonophore (Siphonophora, Physonectae). Bulletin of Marine Science, 76: 699–714.

Dunn, C.W., P.R. Pugh, and S.H.D. Haddock. 2005. Molecular Phylogenetics of the Siphonophora (Cnidaria), with implications for the evolution of functional specialization. Systematic Biology, 54: 916–935.

Haddock, S.H.D., and C.W. Dunn. 2005. The complex world of siphonophores. JMBA Global Marine Environment, 2: 24–25.

Haddock, S.H.D., and J.N. Heine. 2005. Scientific blue-water diving. California Sea Grant.

Haddock, S.H.D., C.W. Dunn, and P.R. Pugh. 2005. A reexamination of siphonophore terminology and morphology, applied to the description of two new prayine species with remarkable bio-optical properties. Journal of the Marine Biological Association of the United Kingdom, 85: 695–707.

Herren, C.M., S.H.D. Haddock, C. Johnson, M.A. Moline, C.M. Orrico, and J.F. Case. 2005. A multi-platform bathyphotometer for fine-scale, coastal bioluminescence research. Limnology and Oceanography: Methods, 3: 247–262.

Miller, S.D, S.H.D. Haddock, C.D. Elvidge, and T.F. Lee. 2005. Detection of a bioluminescent milky sea from space. Proceedings of the National Academy of Sciences, 102: 14181–14184.

Shulman, I., D.J. McGillicuddy Jr., M.A. Moline, S.H.D. Haddock, J.C. Kindle, D. Nechaev, and M.W. Phelps. 2005. Bioluminescence intensity modeling and sampling strategy optimization. Journal of Atmospheric and Oceanic Technology, 22: 1267–1281.